A generation after the events of The Shut Face of Thunder, Thraxis strikes. The anaconda queen resurrects long-dead species of dinosaurs to create an army that allows her to rule the seven continents of Animus like her great ancestors once did. Yet, Thraxis is not concerned only with power. The serpent queen issues a dire warning to the animal world: All animals must unite under her rule before the whale god King Blu attacks and kills them all. Thraxis confronts the whale god, a creature so powerful that none have seen him and survived. Just as the animals fight to overthrow Thraxis, the whale god breaches for the first time in thousands of years. War is inevitable.